Hi Dan,
If I read your question right, then I reckon Vlad has got slightly "the wrong end of the stick" about TTL. What he says is perfectly correct about how film cameras do TTL. However I think you are asking about how Ikelite's digital TTL sensor works.
With the DS50 and DS125, they can be made to flash multiple times to "echo" the output of the internal strobe. The Remote TTL sensor picks up this internal flash, which is often masked to cut out backscatter. The first flashes from that strobe are used for focus and/or metering. The Ikelite strobes using the remote TTL sensor provide the light for this, by firing when the internal fires, and quenching when the internal strobe quenches. When the main flash goes off for the exposure (usually for longer than the metering flash) the strobe again echo this.
Now if I have got this wrong, then please correct me, but I beleive this holds true for setups where the strobes are optically coupled to the internal strobe on the camera.
I also believe that the Sea & Sea DX strobes, when used optically only allow you to ignore the metering flash and fire only on the main flash, rather than providing any intelligence like the Ikelite system, which allows the camera's TTL system to work properly.
As ever TTL is opnly of any real help for exposure in a macro environment, and for wide angle you are best running on manual strobe, manual exposure on the camera.
Sorry this has been sucha rambling post, but I'm trying to get much of this straight in my head also, so corrections are gratefully received.
Cheers
Martyn